Opisthobranchia

Opisthobranchs () is a now informal name for a large and diverse group of specialized complex gastropods which used to be united in the subclass Opisthobranchia. That taxon is no longer considered to represent a monophyletic grouping. Euopisthobranchia is a taxon containing a revised collection of opisthobranchs, and that taxon is considered monophyletic. Euopisthobranchia does not include some "traditional" opisthobranchs such as the Sacoglossa and the Acochlidiacea. The subclass Heterobranchia now contains all the species which used to be assigned to Opisthobranchia, plus all the species in the Pulmonata. The subclass Opisthobranchia included species in the order Cephalaspidea (bubble shells and headshield slugs), the sacoglossans, anaspidean sea hares, pelagic sea angels, sea butterflies, and many families of the Nudibranchia. Opisthobranch means "gills behind" (and to the right) of the heart. In contrast, Prosobranch means gills in front (of the heart). Opisthobranchs are characterized by two pairs of tentacles and a single gill behind and to the right of the heart. With the lack of a heavily mineralized shell, there has been very little fossil record of the group. However, molecular clock studies have suggested that Opisthobranchia emerged as early as the Carboniferous.